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{{for|Spren|more information about spren}}
 
Among many factors that make Roshar unique among other [[Cosmerecosmere]] worlds is the overabundance of [[spren]]. Spren are small pieces of [[Investiture]] that have aquired sentience - and, in some cases, sapience - through the influence of Roshar's living creatures.{{book ref|sa2|3}} They are a mix of different [[Shard]]ic investiture, mainly [[Honor]] and [[Cultivation]], and exist mostly in the [[Cognitive Realm]].{{wob ref|11949}}{{book ref|sa2|1}} However, they can and will often become partially visible in the [[Physical Realm]], and some transition there entirely.
 
Three main groups of spren can be recognized. The nature spren correspond to various natural phenomena, such as [[Spren#Windspren|windspren]] or [[Spren#Flamespren|flamespren]], which appear in the presence of wind and fire respectively. Emotion spren are attracted to emotions, such as [[Spren#Fearspren|fearspren]] appearing when one is afraid or [[Spren#Gloryspren|gloryspren]] for people who feel intense pride.{{book ref|sa2|1}} However, a natural phenomenon or an emotion will not necessarily summon a spren, as they may not be present in the physical place's Cognitive equivalent.
Other than the cremlings, Roshar also features a number of larger crustaceans that are nonetheless smaller than humans. This group includes predatory species, such as the [[axehound]], herbivores like [[axehind]], and near-cremling sized creatures like [[legger]]s and [[larkin]]s, as well as species that can be found on Earth, like crabs.{{book ref|sa3|63}} The larkins in particular are unusual for their flight, as well as the ability to pull Stormlight out of other creatures and absorb it.{{book ref|sa2|i|9}}
 
The largest family of animals on Roshar are known as [[greatshell]]s, which exist both on land and in the seas. All greatshells possess gemhearts and utilize them to attract [[mandra]]s to decrease their own weight. The land greatshells include domesticated herbivores known as [[chull]]s and two large predatory species, the [[chasmfiend]] and the extinct [[lanceryn]]. The marine greatshells include the [[yu-nerig]], which comes on land to pupate and is known to feed on humans,{{book ref|sa1|40}} and the [[Tai-na]], the largest animals known to inhabit the Cosmerecosmere. Their legs are said to be sixty feet long, while the body itself stretches hundreds of feet tall. The back of Tai-na, which sticks out above water, is an ecosystem of its own, inhabited by multiple species of plants and animals, as well as humans.{{book ref|sa2|i|2}}
 
The [[santhid]], a massive marine species with large shell and a number of tendrils, might be considered a greatshell, and houses its own ecosystem, much like the Tai-na; however, as they were considered half-mythical until very recently, it's impossible to state so with any certainty.{{book ref|sa2|1}}
During the lead up to the [[Day of Recreance]], many events occurred which demoralized the Knights Radiant. The [[Sibling]], for some unknown reason, withdrew from [[Urithiru]] and forced the Knights to abandon their ancestral home. Many Radiants blamed this on a perceived unworthiness, including increased conflicts between the [[Windrunners]] and [[Skybreakers]], but scholars of the time disagreed.{{epigraph ref|sa3|70}}{{epigraph ref|sa3|64}} This concern extended to potential infiltration by [[Unmade]].{{epigraph ref|sa3|73}} While this was occurring, the False Desolation was coming to a close. The presumed success in imprisoning Ba-Ado-Mishram leading to the singers having their Connection and Identity ripped out of them likely influenced the Radiants.{{book ref|sa3|17}} In the midst of this turmoil, the Radiants discovered the truth about the origins of humans on Roshar. The singers had been the original inhabitants of Roshar. The humans came as refugees from their [[Ashyn|homeworld]], which they had destroyed with surges. After being taken in by the singers and given [[Shinovar]], they invaded Roshar and started the Desolations. While Honor had comforted previous generations who learned this, this time he was dying. He raved about the [[Dawnshard]]s and promised the Radiants they would destroy Roshar like they had their homeworld.{{book ref|sa3|113}}
 
As a result of these revelations, the nine of the ten orders decided to abandon their oaths and kill their spren. They believed that this would prevent the destruction of Roshar Honor warned them about.{{book ref|sa3|113}} At the locations they left their [[Shardplate|platePlate]] and [[Shardblade|bladesBlades]], men quickly began fighting over the weapons, leading to much death and destruction.{{book ref|sa1|52}} The Radiants, other than the Skybreakers, were quickly turned on and killed because of the sense of betrayal felt.{{epigraph ref|sa2|41}} Society for both spren and humans drastically changed in the aftermath. Those with Shardblades quickly consolidated their power, including using ''[[Arts and Majesty]]'' to justify women not wielding them.{{wob ref|6245}} It is likely that this is when [[lighteyes]] truly came to power. The genocide decimated the populations of spren societies. Most of the survivors were children forced to spend centuries without nurture or guidance. Some societies took two thousand years to fully recover. {{book ref|sa3|47}}
 
==== The Hierocracy and the War of Loss ====
:''Main articles:'' [[Hierocracy]] and [[War of Loss]]
The [[Hierocracy]] began when the [[Vorinism|Vorin Church]] decided to conquer the world and run the government rather than just provide spiritual guidance.{{book ref|sa1|36}} They claimed their conquering was for the good of the people. Some of the most powerful ardents claimed to have visions from the [[Almighty]], which they used to solidify their rule and obfuscate their religion. One of their main goals was to destroy knowledge of the failures of Vorinism, including [[Shadesmar]], the Knights Radiant, and the Recreance.{{book ref|sa2|3}}{{book ref|sa1|45}} They were primarily successful, as even the major libraries on Roshar are missing this information. Before the Hierocracy could spread to the west, [[Sadees]], the Sunmaker, united the princedoms of [[Alethkar]] to drive out the HIerocracyHierocracy.{{book ref|sa1|18}}{{book ref|sa1|24}} They fought in many epic battles the likes of which have not been seen since in what came to be known as the [[War of Loss]].{{book ref|sa1|52}} The Sunmaker successfully cast down the Hierocracy and stated the visions were a fraud.
 
Sadees's win in the War of Loss affected much of eastern Roshar. Sadees declared himself king of Alethkar and began a conquest of Roshar.{{book ref|sa3|34}} Although the extent of his empire is not exactly known, [[Herdaz]] and [[Azir]] were both included. He committed genocide in Azir and killed ten percent of the population.{{book ref|sa3|42}} His empire quickly collapsed after his death, however, as he did not choose an heir, and his ten sons divided the kingdom into princedoms and became [[highprince]]s.{{book ref|sa3|19}} The effects of the Hierocracy on Vorinism can still be felt in the modern time. The central leadership of the church was destroyed and split into mostly independent [[Devotaries]].{{book ref|sa1|33}}{{book ref|sa1|36}} The ardents became slaves who could not own land or property.{{book ref|sa1|45}}{{book ref|sa1|59}} Their revisions to historical texts also perpetuate, and it is difficult to find information about pre-Hierocracy history or the Knights Radiant.{{book ref|sa2|3}}
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