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Some species of native Rosharan flora are edible for humans. This includes a number of fruits as well as a rice-like plant called [[tallew]], and [[lavis]], a type of rockbud that produces grain. Tallew and lavis appear to be the major ingredients of Rosharan diet, with massive farmlands devoted to them throughout the continent.{{book ref|sa1|4}} In the West, the same niche appears to be filled by [[clema]].{{book ref|Edgedancer|6}}
 
Additionally, there exist a number of plants with known healing properties. This includes antiseptics, such as the sap of [[knobweed]] and [[fourleaf]], as well as [[lister's oil]]; treatments for specific diseases, such as [[bitterleaf]] for epilepsy and [[bloodivy]] for the [[grindings]]; anesthetics like [[winterwort]] and [[firemoss]]; and other, more specific medicines, such as [[denocax salve|denocax]], which encourages skin healing.
 
By contrast, several species of plant are noted to have detrimental effect on humans. Firemoss is known to be highly addictive, and [[blackbane]]'s leaves, once dried, are extremely poisonous. Eating shells of rockbuds is similarly known to be unhealthy, although the [[Unkalaki]] are capable of eating them without problems, likely thanks to their [[singer]] ancestry.{{wob ref|930}} While not all plants are edible, some, like [[breachtree]] and [[narbin]], are grown to make fabrics.{{book ref|sa1|25}}{{book ref|sa1|45}}
Multiple native species, as well as at least one imported animal, have formed symbiotic relationships with [[spren]]. This appears to be faciliated by [[gemheart]]s, crystals that grow naturally within the bodies of those creatures.
 
* Larger animals on Roshar utilize bonds with [[Mandramandra]]s, or luckspren, to decrease the pull of gravity on them, allowing creatures such as [[Chasmfiendchasmfiend]]s to exist without collapsing upon themselves. In the case of [[Skyeelskyeel]]s, the same bond, coupled with smaller size, permits them to fly.
* The native sentient species, the [[Singer]]s, bond with a variety of spren to attain a number of forms, which alter their mental state, biological functions and appearance. While bonded, the spren remain within the gemheart.
* [[Ryshadium]]s bond with what appears to be musicspren, attaining higher intelligence and physical strength. It is, however, unclear so far whether or not they have gemhearts.{{au ref|rosharan}}
 
;Non-shelled animals
Mammals dwell mostly in Shinovar, though a few species have been successfully exported Easteast. Those include horses, which are widely used as far east as Alethkar, and hogs, which are the primary source of non-soulcastSoulcast meat. While most mammals are domesticated, hogs, as well as minks and rats,{{book ref|sa1|3}} can be found in the wild. Several species of birds are known to exist, although they are all known as "chickens" among Rosharans. This includes actual chickens as well as more colourful birds, such as parrots. In contrast to mammals, they do not seem to do well in the wild, as all examples observed so far are domesticated. Amphibians, such as the [[lurg]] can also be found, as well as wide variety of fish, including both salt- and seawater ones. Cephalopods also exist, albeit with unusual properties - [[zatalef]], for example, is described as bat-like, implying the ability to fly.{{file ref|Alethi_Glyphs_Page_1.jpg}} A category without a clear Earth analogue are [[sarpenthyn]], which appear to be similar to both mollusks and cephalopods, albeit without shells and with smaller number of arms.{{book ref|sa2|55}}
 
;Shelled animals
The largest known group of animals on Roshar are crustaceans. Those can be of various sizes, from finger-long to four-stories-high. They are all noted as having purpleviolet blood and multiple stages of life, with the initial stage being called a larva and the process of switching between stages called pupation.{{file ref|Chull_Life_Cycle.jpeg|Chull Life Cycle}} The creation of shells is aided by the presence of crem in the rain, which the creatures can process into minerals.
 
One of the largest crustacean family are [[cremling]]s, a group of tiny, multi-legged scavengers preying mostly on deceased animals, although they will also feed on uncovered plants after highstorms. Cremlings come in variety of colours, with different features, numbers of legs, and behaviours. While most are individuals, some are part of the [[Sleepless]] swarms, and as such share a sentient hivemind. The [[Sleepless]] cremlings are bred for particular purposes, such as imitating human body parts.
Other than the cremlings, Roshar also features a number of larger crustaceans that are nonetheless smaller than humans. This group includes predatory species, such as the [[axehound]], herbivores like [[axehind]], and near-cremling sized creatures like [[legger]]s and [[larkin]]s, as well as species that can be found on Earth, like crabs.{{book ref|sa3|63}} The larkins in particular are unusual for their flight, as well as the ability to pull Stormlight out of other creatures and absorb it.{{book ref|sa2|i|9}}
 
The largest family of animals on Roshar are known as [[greatshell]]s, which exist both on land and in the seas. All greatshells possess gemhearts and utilize them to attract [[Mandramandra]]s to decrease their own weight. The land greatshells include domesticated herbivores known as [[chull]]s and two large predatory species, the [[chasmfiend]] and the extinct [[lanceryn]]. The marine greatshells include the [[yu-nerig]], which comes on land to pupate and is known to feed on humans,{{book ref|sa1|40}} and the [[Tai-na]], the largest animals known to inhabit the Cosmere. Their legs are said to be sixty feet long, while the body itself stretches hundreds of feet tall. The back of Tai-na, which sticks out above water, is an ecosystem of its own, inhabited by multiple species of plants and animals, as well as humans.{{book ref|sa2|i|2}}
 
The [[aanthidsanthid]], a massive marine species with large shell and a number of tendrils, might be considered a greatshell, and houses its own ecosystem, much like the Tai-na; however, as they were considered half-mythical until very recently, it's impossible to state so with any certainty.{{book ref|sa2|1}}
 
;Domestication
=== The Era of Solitude ===
{{quote
|The Voidbringers were cast back into the Tranquiline Halls. The Heralds followed to force them out of heaven as well, and Roshar’s Heraldic Epochs ended. Mankind entered the Era of Solitude., The modernthe era in which the [[Stormlight Archive]] is set.
|[[Kabsal]] to [[Shallan]]{{book ref|sa1|45}}
}}
During the lead up to the [[Day of Recreance]], many events occurred which demoralized the Knights Radiant. The [[Sibling]], for some unknown reason, withdrew from [[Urithiru]] and forced the Knights to abandon their ancestral home. Many Radiants blamed this on a perceived unworthiness, including increased conflicts between the [[Windrunners]] and [[Skybreakers]], but scholars of the time disagreed.{{epigraph ref|sa3|70}}{{epigraph ref|sa3|64}} This concern extended to potential infiltration by [[Unmade]].{{epigraph ref|sa3|73}} While this was occurring, the False Desolation was coming to a close. The presumed success in imprisoning Ba-Ado-Mishram leading to the singers having their Connection and Identity ripped out of them likely influenced the Radiants.{{book ref|sa3|17}} In the midst of this turmoil, the Radiants discovered the truth about the origins of humans on Roshar. The singers had been the original inhabitants of Roshar. The humans came as refugees from their [[Ashyn|homeworld]], which they had destroyed with surges. After being taken in by the singers and given [[Shinovar]], they invaded Roshar and started the Desolations. While Honor had comforted previous generations who learned this, this time he was dying. He raved about the [[Dawnshard]]s and promised the Radiants they would destroy Roshar like they had their homeworld.{{book ref|sa3|113}}
 
As a result of these revelations, the nine of the ten orders decided to abandon their oaths and kill their spren. They believed that this would prevent the destruction of Roshar Honor warned them about.{{book ref|sa3|113}} At the locations they left their [[shardplateShardplate|plate]] and [[shardbladeShardblades|blades]], men quickly began fighting over the weapons, leading to much death and destruction.{{book ref|sa1|52}} The Radiants, other than the Skybreakers, were quickly turned on and killed because of the sense of betrayal felt.{{epigraph ref|sa2|41}} Society for both spren and humans drastically changed in the aftermath. Those with shardbladesShardblades quickly consolidated their power, including using [[Arts and Majesty]] to justify women not wielding them.{{wob ref|6245}} It is likely that this is when [[lighteyes]] truly came to power. The genocide decimated the populations of spren societies. Most of the survivors were children forced to spend centuries without nurture or guidance. Some societies took two thousand years to fully recover. {{book ref|sa3|47}}
 
==== The Hierocracy and the War of Loss ====
:''Main Article: [[Spheres]]''
 
The primary form of currency found on Roshar are spheres, pieces of gemstone encased in a sphere of glass with one side flattened to prevent rolling.{{book ref|sa1|27}} The gemstones can be any of the ten [[Polestone]]s and there are three denominations: chip, mark, and broam. The ten gemstones are organized into five value levels with some gem types having equivalent values.{{wob ref|10402}} These value levels are determined by [[Soulcasting]] properties, so diamonds, which can create quartz, crystal, and glass, have the lowest value, and emeralds, which can create grain, have the highest.{{book ref|sa1|3}} Spheres, like all gems on Roshar, have the ability to hold and leak stormlightStormlight. Spheres with light are referred to as infused and those without are referred to as dun. Although dun spheres can be used as currency, people are suspicious of them because light is the easiest way to determine a sphere is real.{{book ref|sa1|17}}
 
Because of their ability to hold stormlightStormlight, spheres are commonly used on Roshar as a source of light. The light from spheres is considered to be steadier and more consistent than traditional light sources such as candles.{{book ref|sa1|23}} It can be expensive, however, as spheres are also used for currency, so the use of spheres as a source of light is often a sign of wealth. Diamonds are considered the best light source because it is both the cheapest and the closest to natural light.{{book ref|sa1|8}}
 
The denomination of spheres in eastern Roshar is weaker than in the west, with some people drilling holes in spheres to carry them on strings or using chunks of gemstone instead.{{book ref|sa2.5|6}}
Within [[Shadesmar]], the sentient [[spren]] have their own countries.{{book ref|sa2|1}} Among those is a [[honorspren]] state, with its capital in the city of [[Lasting Integrity]] and a number of strongholds, as well as a [[Highspren]] country, and a city-state of the [[Cryptic]]s.{{book ref|sa3|102}}{{book ref|sa2|epilogue}}{{book ref|sa2|3}} There are also free cities, such as [[Celebrant]], where various spren intermingle. This includes [[Cultivationspren]], which, though ruled by a council called the [[Ring]], do not have holdings of their own.{{book ref|sa3|102}}{{book ref|sa2.5|2}}{{book ref|sa2.5|1}} However, even the "ethnic" spren states do not appear to be strictly isolationist - for example, honorspren military ships are partially crewed by [[Spren#Mistspren|mistspren]].{{book ref|sa3|108}}
 
As the inversion of the [[Physical Realm]] and its supercontinent, Shadesmar is mostly composed of the sea of beads, with a number of narrow peninsulas and islands corresponding to Physical rivers and lakes.{{map ref|sea of lost lights}} As such, travel is done mostly by boat. Merchant and military ships, pulled by teams of harnessed [[Mandramandra]]s, traverse the vast spaces between the cities, ferrying goods, news and travellers.{{book ref|sa3|99}} Trade with lands outside of Roshar isn't unusual - in Celebrant, one may buy canned food (presumably from [[Scadrial]] or some [[worldhopper]] settlement) and [[Nalthis |Nalthian]] artwork. The price of various items is largely based on their permanence in the Cognitive Realm, with the most persistent objects selling for thousands of broams.{{book ref|sa3|102}}
 
The relationship between various spren kinds appears to have its own share of political turmoil. Several years prior to the [[Everstorm]], the honorspren tried and failed to conquer the rest of Shadesmar, making the other spren unfriendly towards them. By contrast, Cryptics are generally understood to be unwilling to expand beyond their city; however, they are also widely considered to be uncomfortable company.{{book ref|sa3|47}}
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